Following his discharge, Madison struggled to adjust to civilian life and was eventually admitted to the Clinic for Socially Maladjusted Super-Beings. There he met Roger Bochs, a brilliant but physically disabled inventor. The two formed a close friendship, bonding over their shared sense of alienation. Their resistance to conventional therapy drew the attention of James MacDonald Hudson, who recruited them into Gamma Flight, the training division of Alpha Flight. Both men later advanced to Beta Flight.

When Department H was temporarily shut down, many former members turned to crime or independent activities. Jeffries and Bochs, however, remained committed to using their abilities constructively. Bochs developed the Box armor to compensate for his physical limitations, while Jeffries assisted in its design and maintenance, becoming deeply familiar with its systems.

During the rise of Omega Flight under Jerome Jaxon, Bochs infiltrated the organization but was discovered. The resulting confrontation led to the deaths of both Jaxon and Guardian. In the aftermath, Alpha Flight reorganized, and Jeffries was brought into the team as both a field operative and technical specialist.

Jeffries quickly proved invaluable, particularly in the reconstruction and modification of Guardian’s armor for Heather Hudson, enabling her to assume the identity of Vindicator. During subsequent missions, Jeffries also played a key role in defeating the robotic duplicate Delphine Courtney, further cementing his place within the team.

Tragedy struck when Roger Bochs’ physical and mental condition deteriorated. After transferring his consciousness into the Box armor permanently, Bochs became increasingly unstable. In a desperate attempt to stabilize himself, Bochs merged with Lionel Jeffries, creating the immensely powerful entity known as Omega. The combined being proved uncontrollable, forcing Madison into a final confrontation. With great reluctance, Jeffries destroyed Omega, resulting in the deaths of both his brother and Bochs.

In the aftermath of this loss, Jeffries assumed control of the Box armor himself, becoming Box (IV). Unlike Bochs, Jeffries could directly interface with the armor using his mutant abilities, allowing for far greater adaptability and power. As Box (IV), he became one of Alpha Flight’s most powerful and versatile members.

Over time, Jeffries’ role within Alpha Flight expanded beyond combat. He became the team’s primary engineer and technological specialist, continuously upgrading the Box armor and supporting the team’s operations. His relationship with Heather Hudson grew increasingly close, providing him with a measure of emotional stability following the trauma of his past.

Jeffries has remained a consistent presence within Alpha Flight through its various incarnations, including its later government-sanctioned operations. Despite recurring personal struggles and the lingering impact of his brother’s fate, he has continued to serve as a hero, using his powers and intellect to protect both Canada and the wider world.

In more recent years, Jeffries has demonstrated an even greater mastery of his abilities, blurring the line between man and machine as he integrates more deeply with the Box armor. His powers have evolved to the point where he can reshape complex systems in real time, making him one of the most formidable technopath-adjacent mutants in existence.

Jeffries’ powers allow him to effectively “think machinery into new forms,” making him uniquely suited to operate and continuously upgrade the Box armor. His control is precise enough to function at microscopic levels, enabling advanced structural modifications in real time.

Limitations: Jeffries’ powers are limited to non-organic materials such as metal, plastic, and glass. His effectiveness depends on available material and his mental state.